On leaves, small brown spots appear first. These enlarge, develop a lighter-coloured central area, and become irregular or angular in shape.
On stems, elongated dark-brown lesions appear, usually near leaf axils or where stems have rubbed against the supporting wire. Infection spreads from these points and whenever the stem becomes completely girdled the shoot suddently wilts and fruits collapse.
On fruit, spots first appear as pinpricks, which enlarge into sunken circular lesions with brownish centres. Eventually the rind round the diseased area becomes wrinkled and the fruits shrivel and drop.
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