Yellow spots without a definite margin appear on the upper leaf surface, and olive-green to brown or purplish masses of spores and velvety growth appear on the lower leaf surface. Disease symptoms appear first on the older leaves. As the disease progresses, the leaves become chlorotic, then necrotic, followed by drying and defoliation.
Although this is primarily a foliar disease, unripe fruit may occasionally develop dark leathery lesions.
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